Frost damages plants when ice crystals develop in the cells, denying water to plant tissue and impeding the flow of hydration. "Frost-damaged leaves or twigs appear water-soaked, wither, and turn a dark brown or black," reports the University of California Division of Agriculture. Young or regrafted fruit trees are particularly vulnerable.
The temperature at which frost will damage tree fruits like apples and pears, and stone fruits such as apricots and peaches, is largely dependent on the variety and species. The University of Arizona's website recommends wrapping the trunk of young citrus and other sensitive trees with weatherproof paper or cloth.
Commercial farmers employ a variety of techniques---like fans---to protect their crops from frost damage. Michigan State University Extension Service advises growers to remove obstructions to help cold air more quickly escape from the growing area. "There are numerous sites where a degree or two could be gained by removing obstructions from air drainage passages under radiation freeze conditions," the extension service says.